About the PhD

a highly multidisciplinary training for the study of environmental complexity

The course aims at fostering the interdisciplinary integration of Natural Sciences, including Physics, Biology and Earth Sciences, with Veterinary and Agricultural Sciences, Medicine and Pharmacology. PhD students will be trained in the study of past and present biotic and abiotic processes that shaped our environment, taking into consideration past and extant interactions and functional relationships, while making full use of ecological, evolutionary, geological and geomorphological approaches.

PhD students will receive specific training in broadly diverse field and laboratory techniques. These will range from geological and ecological sampling, to advanced microscopy techniques and biochemical and molecular assays. They will also receive training in Geographic Information Systems (GIS), statistical modelling and bioinformatics.

Head of the PhD course
Prof. Nicola Saino (nicola.saino@unimi.it)


Restricted PhD Course board (giunta)


Broadly, areas of interest include the following



MARCH 18th, 2019, h. 2:00 p.m.

Room 108DX, Via Celoria 26, Milano



Final exam - XXXI Cycle

FEBRUARY 5th, 2019, h. 10:30

Room BS, Via Celoria 26, Milano



Marco Bonelli added a new publication (10/03/2019)

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Magda Zanzottera added a new publication (10/03/2019)

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    March 22, 2019

  • Seminar by Prof. Jana Kusch, entitled: Cyclic nucleotide-dependent gating in HCN pacemaker channels
  • March 26, 2019

  • Seminar by Prof. Matthew TUCKER, entitled: Molecular and compositional features of female germline development in flowering plants
  • March 29, 2019

  • Seminar by Dr. Filippo DEL BENE, entitled: Regulation of retinal progenitor cell differentiation by redox signaling and disease modeling in zebrafish with genomic specific base editors
  • April 16, 2019

  • Seminar by Jack Tuszynski, entitled: 100 Years from Quantum Physics to Quantum Biology . How Long to Quantum Medicine?